You should always type a SUBJECT in the "subject" line...
Anyway, the Mark I should be modifiable for 3-500Zs, as I did this many (many!) years ago on an old Mark-VI, which is the six meter version of the same amp. It originally used 3-400Zs also, and of course within about one year, Eimac discontinued those (like in about 1969 or something).
My mod is a hazy memory now, but I recall mounting the tube sockets down lower in the chassis, using standoffs to drop the sockets down, and that created enough clearance for the 500Zs to fit just fine without changing anything else.
The original cooling fan seemed to blow enough air in the right places that it didn't require any changes.
I added a few volts of cathode bias with a 50W or 100W stud zener, as the idling current with the 500Zs was a bit high until a bit of positive cathode bias was added to bring it down some. I think in the end, I had about 100mA idling current for the pair of tubes.
This was not a major modification, and I'm pretty sure the Mark I sheet metal is the same design, maybe even the same metal, as the Mark-VI.
Obviously, a sloppy job can be done anytime one wishes to be sloppy! But mine came out fine, and I sold the amp for a very premium price years later.
WB2WIK/6