James,
You might consider my comment it was 100% your fault harsh, but it is true. I'm not one to dance around the issue. :-)
It take terrible heat to actually melt glass. This is NOT directly related to fuse size or drive power, it is related to tuning and operation. Both are under the direct control of the operator.
The dissipation or heat produced is a direct function of the input power to the tubes, and the output power to the load. If you had 2300 volts at 500 mA plate current, that would be 2300*.500 = 1150 watts of power. If the real power going into the feedline was 740 watts, that would be 1150-740 = 410 watts of heat.
Each tube in your amplifier is rated at 160 watts of steady anode heat, or 320 watts total. On CW or voice, because the load is varying and not steady, 410 watts is OK on peaks. The thermal averaging in the mass of the tube will allow heat peaks well over 320 watts. This is how the amp should have run.
On SSB voice, you must have operated for an extended time with much more than 500 watts of average heat. That is what it would take to suck the glass in on a 572B tube. This means the peak heat must have been terrible!
This might make your head swim, but the concept is the important thing.
Someone else suggested bad components or parasitics, but in truth that is very unlikely to be a root cause. Even if it were true, watching the power output meter and the plate current would have warned you right away.
You need a good output meter and need to know how to use it, and you also need to understand how to read the plate current meter and what it means. Otherwise you will be right back into expensive trouble.
I'll add a plate current vs. power output table on my web site to help you and others. I'll work on that today. Your problem, and the responses here, have caused me to realize something lacking in amplifier instructions. Since I design amplifiers and write manuals, I can do a better job because of reading about your problem and the suggestions of others. You spent the money, but I learned something that is missing from manuals. Every manual should have a plate current vs. power output chart or graph!!
I've never thought of this before but I can see how a plate current vs power output graph or chart would be one of the most valuable assets of all, other than telling people to watch grid current.
73 Tom